MALACOLOGIA
http://www.instituteofmalacology.org
One of the World’s Leading Malacological Journals
Journal Impact Factor 13.5 (2019)
Founded in 1961
OFFICERS
Editor-in-Chief CHARLES LYDEARD
Editorial Office
CHARLES LYDEARD
Morehead State University
Department of Biology and Chemistry
103 Lappin Hall
Morehead, Kentucky 40351, U.S.A.
clydeardmalac@gmail.com
Composition Editor
CLAUDIA WILKE
Giessen, Germany
claudiawilke@hotmail.com
Honorary Associate Editor
JOHN B. BURCH
Honorary Associate Editor
GEORGE M. DAVIS
Business Manager
DAVID CAMPBELL
110 S Main St, Box 7236
Boiling Springs, North Carolina 28017, U.S.A
malacologia@gardner-webb.edu
Copy Editor
EUGENE COAN
Santa Barbara Museum of Natural History
Santa Barbara, California, U.S.A.
genecoan@gmail.com
Advisory Editor
THOMAS WILKE
Justus Liebig University
Giessen, Germany
tom.wilke@allzool.bio.uni-giessen.de
MALACOLOGIA is published by the INSTITUTE OF MALACOLOGY, U.S.A.
BOARD OF DIRECTORS
President
ELLEN STRONG
National Museum of Natural History
Washington D.C.
Vice President
RÜDIGER BIELER
Field Museum of Natural History, Chicago, Illinois
Treasurer
CHARLES LYDEARD
Morehead State University
Morehead, Kentucky
Secretary
PAULA M. MIKKELSEN
Field Museum of Natural History, Chicago, Illinois
Members
RÜDIGER BIELER
Field Museum of Natural History Chicago, Illinois
JOHN B. BURCH
University of Michigan, Ann Arbor, Michigan
CAROLE S. HICKMAN
University of California, Berkeley, California
ALAN KOHN
University of Washington, Seattle, Washington
DIARMAID Ó FOIGHIL
University of Michigan, Ann Arbor,
Michigan
Copyright © 2020 by the Institute of Malacology
ISSN: 0076-2997 (print)
ISSN: 2168-9075 (Online)