MALACOLOGIA
http://www.instituteofmalacology.org
One of the World’s Leading Malacological Journals
Journal Impact Factor 13.5 (2019)
Founded in 1961
OFFICERS
Editor-in-Chief
REBECCA J. RUNDELL
State University of New York
Syracuse, New York USA
Malacologia E-mail Address: rundellmollusca@gmail.com
Editorial Office
REBECCA J. RUNDELL
State University of New York
Syracuse, New York USA
Malacologia E-mail Address: rundellmollusca@gmail.com
Business Manager
DAVID CAMPBELL
110 S Main St, Box 7236
Boiling Springs, North Carolina, U.S.A., 28017,l
malacologia@gardner-webb.edu
Copy Editor
EUGENE COAN
Santa Barbara Museum of Natural History
Santa Barbara, California, U.S.A.
genecoan@gmail.com
Production Editor
PAULA M. MIKKELSEN
Field Museum of Natural History
Auburn, Maine, U.S.A.
MALACOLOGIA is published by the INSTITUTE OF MALACOLOGY, U.S.A.
BOARD OF DIRECTORS
President
ELLEN STRONG
National Museum of Natural History
Washington D.C.
Vice President
RÜDIGER BIELER
Field Museum of Natural History, Chicago, Illinois
Treasurer
CHARLES LYDEARD
Morehead State University
Morehead, Kentucky
Secretary
PAULA M. MIKKELSEN
Field Museum of Natural History, Chicago, Illinois
Members
DIARMAID Ó FOIGHIL
University of Michigan, Ann Arbor,
Michigan
CAROLE S. HICKMAN
University of California, Berkeley, California
ELIZABETH SHEA
Delaware Museum of Natural History
Wilmington, Delaware
GUSTAV PAULAY
Florida Museum of Natural History
Gainesville, Florida
Copyright © 2023 by the Institute of Malacology
ISSN: 0076-2997 (print)
ISSN: 2168-9075 (Online)